If the Five Elders’ Yokai ability works similarly to Devil Fruits, then his Yami Yami no Mi is going to be key to getting around their regeneration. On top of that, he has the power of the Gura ...

This is why they have Yokai based names instead of having Devil Fruits that allow them to transform into the said creatures. The Five Elders are magnificent in terms of what they offer as villains.